"I have a longstanding interest in energy metabolism and responses of animals, particularly agriculturally important species, to the environment. We have also investigated how body control systems respond when two or more controls are in conflict. For example, when exercise induces a heat load in panting animals, the thermoregulatory system seeks to improve heat loss by increasing ventilation. This response results in a precipitous drop in arterial CO₂ tension, which opposes the increase in ventilation. How do these control systems resolve this conflict.

More recently, I have developed a strong interest in student learning. Some questions of interest include:

1. Do simplifications of physiological phenomena introduce errors in student thinking?
2. What is it, specifically, about computer-based education that makes it a valuable learning tool?
3. What do our exams actually measure: knowledge, IQ, or something else?
4. How do professional physiologists' and students' reasoning processes differ?"